The LEDs in this circuit produce a chasing pattern similar the running LEDs display in video shops. In fact the effect is called: "Running Hole." All transistors will try to come on at the same time when the power is applied, but some will be faster due to their internal characteristics and some will get a different turn-on current due to the exact value of the 22u electrolytics. The last 22u will delay the voltage-rise to the base of the first transistor and make the circuit start reliably. It is very difficult to see where the hole starts and that's why you should build the circuit and investigate it yourself. The circuit can be extended to any number of odd stages.